College of Sciences staff by location (campus)

Contact details +6469517835

Prof Mary Morgan-Richards PhD

Professor in Evolutionary Biology

Doctoral Supervisor
School of Food Technology and Natural Sciences

I am an academic at Massey University, teaching biodiversity, conservation and evolution. I love working with New Zealand invertebrates (weta, stick insects, snails) and am particularly keen on hybrid zones. Hybrid zone are where distinct populations meet and produce hybrids, such zones are stable because the hybrids are less successful than their parents. Hybrid zones provide a window into the process of speciation, gene flow and capture of adaptive alleles. I study the areas where different chromosome races of the tree weta Hemideina thoracica met and mate and produce hybrid offspring. By comparing the width of different frequency clines one can measure relative fitness disadvantage suffered by the hybrids. New Zealand geckos also form hybrid zones where large geckos meet small geckos. My students and I use genetic tools to study the population genetics of skinks and geckos and inform conservation management decisions. Collaborations with paleontologists have lead my students and I to investigate the role of cladogenesis in geologically rapid changes in morphology. This work uses New Zealand’s excellent marine fossil record and extant marine gastropods.

Before coming to Massey University I did post-docs in the UK (University of St Andrews and The Natural History Museum London) and NZ (Otago and Canterbury Universities). I currently work 0.7 of a full time position as an academic and juggle my family life.

I am an evolutionary biologist working within the Ecology group (SNS) on the Manawatu campus. My research expertise is in hybridisation of genetically distinct populations or species, and the process of speciation. I teach population genetics, conservation biology, and New Zealand Natural History to undergraduate students. My graduate students use native invertebrates such as weta, stick insects, marine snails and feather lice to test evolutionary hypothesis about the formation of new species and the distribution of genetic and adaptative diversity

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Professional

Contact details

Ph: 84835
Location: 1.17, AgHort
Campus: Manawatu
Ph: 85731
Location: D5.25 (farside lab), ScienceTower D
Campus: Turitea

Qualifications

Doctor of Philosophy - Victoria University (1995)

Certifications and Registrations

Licence, Supervisor, Massey University

Research Expertise

Research Interests

My graduate students and postdoctoral fellows work on a range of native species focusing on evolutionary ecology, speciation and phylogeography. We use New Zealand invertebrates such as weta, stick insects and snails to address evolutionary questions and study species interactions in natural ecosystems. My students and I study weta and gecko hybrid zones. I collaborate with paleontologists (GNS Science) to investigate evolutionary relationships of extant molluscs and use their excellent fossil record to estimate rates of DNA evolution, patterns of extinction and speciation and the problem of convergence.

Research Opportunities

Biodiversity (01/03/2013) Opportunities exist within my research group (evolves@massey.ac.nz) to investigate local biodiversity using the cave weta as a model system. Many species await discovery and time is short for many forest habtats. This project will use systematic tools to document levels and patterns of New Zealand forest biodiversity
hybrid zones (01/03/2013) Opportunities exist within my research group (evolves@massey.ac.nz) to investigate the level of hybridisation within and among tree weta species. Using chromosome and molecular genetic tools gene flow will be measured and evidence of reproductive isolation sought.
conservation genetics (01/03/2013) Opportunities exist within my research group (evolves@massey.ac.nz) to investigate questions relating to biodiversity, conservation genetics and molecular ecology. For example work on New Zealand endemic geckos is needed to ensure management decisions conserve our current species-level diversity. This research would require detailed morphological and genetic study of gecko populations to delineate species boundaries and would be carried out in collaboration with Department of Conservation staff.
Ecology (01/07/2022) An ecologically friendly solution is required to manage an endemic cricket (wētā) in New Zealand vineyards. The factors that determine the distribution and abundance of this species will be studied

Thematics

21st Century Citizenship, Health and Well-being

Area of Expertise

Field of research codes
Animal Systematics and Taxonomy (060301): Biogeography and Phylogeography (060302): Biological Adaptation (060303): Biological Sciences (060000):
Earth Sciences (040000):
Evolutionary Biology (060300): Evolutionary Impacts of Climate Change (060306): Genetics (060400): Host-Parasite Interactions (060307): Phylogeny and Comparative Analysis (060309): Population, Ecological and Evolutionary Genetics (060411): Speciation and Extinction (060311): Zoology (060800)

Keywords

Evolution, Speciation, hybridisation, gene flow, hybrid zones, molecular genetics, cytogenetics, analytical methods of population genetics, Genomics, New Zealand Orthoptera, Stick Insects, Chromosomes, DNA content.

Research Projects

Summary of Research Projects

Position	Current	Completed
Project Leader	2	22

Completed Projects

Project Title: Punctuated evolution: is rapid morphological change linked to speciation?

The study of fossils provides an impression of morphological evolution made up of long periods of constrained evolution when nothing changes interspersed with geologically sudden leaps in form. Many interpret these abrupt changes in morphology as being the result of speciation (punctuated equilibrium). Other explanations for this pattern are possible and include rapid adaptation without speciation, the invasion of species from elsewhere, or hybridisation. New Zealand has one of the world's best fossil records for marine snails and many lineages in the rocks have relatives alive today. For the first time we can study the morphology and the molecular evolution of the same snail lineages united to produce a time-space integrated view of phenotypic divergence. This project brings together a unique combination of paleontologists, molecular geneticists and phylogeographers to study speciation in the past and present. Using mathematical analysis of shell shape changes and the latest DNA sequencing tools we will determine whether the timing of morphological change coincides with speciation inferred from molecular phylogenetics. By answering the fundamental question 'Is morphological change the result of species formation'. We will place New Zealand at the forefront of speciation research internationally.

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Date Range: 2013 - 2016

Funding Body: Marsden Fund - Full

Project Team:

Prof Mary Morgan-Richards - Project Leader
Prof Steven Trewick - Team Member

Project Title: Can weta coexist?

If two species compete for the same resources such as food and shelter one species may exclude the other from a region. In cases of coexistence we expect to see divergent selection to result in ecological differences among species. Tree weta have only narrow regions of sympatry that may move depending on the climate and their competitive interactions. We are investigating the diet and behavior of two tree weta where they occur together in the Manawatu. By considering assortative mating, gene flow, optimal nutrient targets, morphology and diet we can assess the likelihood that the two species can coexist long term.

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Date Range: 2013 - 2013

Funding Body: Massey University

Project Team:

Prof Mary Morgan-Richards - Project Leader

Project Title: What limits a weta?

The distribution of a species is controlled by a combination of environmental factors and biotic interactions. For example temperature must be warm enough to allow growth, and the right foods must be available in the environment. We investigated the factors limiting the expansion of the range of the common Wellington tree weta species Hemideina crassidens. We inferred past range changes from patterns of genetic diversity over its range, and examined the competitive interactions that might prevent sympatry with the Auckland tree weta. We are investigating adaptation to high and low elevation sites that might allow the species to expand its range and the potential role of gene flow in limiting adaptation.

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Date Range: 2010 - 2013

Funding Body: Massey University

Project Team:

Prof Mary Morgan-Richards - Project Leader
Prof Steven Trewick - Project Leader

Project Title: TTP Fellowship - Priscilla McAllum-Wehi - Invertebrate indicators of ecological change? Weta, putaputaweta and forest health

Conservation biology now recognises that understanding the functional relationships among organisms is essential for ecosystem health. Just as knowledge of whakapapa is complex and non-linear, optimal ecosystem functioning also relies on understanding linkages between species. Hence the contribution of species such as w?t? to ecosystem services such as seed dispersal, and understanding their trophic position in food webs, helps define these linkages. This project will build on M?ori knowledge of the relationships between culturally iconic invertebrates (w?t? and pepetuna (the caterpillars of the p?riri moth) and trees (putaputaweta and k?tukutuku). By focusing on these relationships, we emphasise (a) the importance of species relationships within forest ecosystems in achieving ecosystem health and sustainability and (b) contribute to a better understanding of the role of these taonga species. We will examine the complex set of interactions between these species using stable isotope and molecular analyses in conjunction with experimental ecological studies to elucidate food web pathways and ecosystem services that connect to w?t?, as well as likely costs and benefits to the tree species, such as putaputaweta, that host them.

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Date Range: 2009 - 2012

Funding Body: Foundation for Research, Science & Technology

Project Team:

Prof Mary Morgan-Richards - Project Leader
Dr Priscilla McAllum - Team Member

Project Title: Alpine plant-insect mutualisms

Many plant species reply on insects to transfer pollen between the flowers of different plants to ensure outcrossing and successful offspring. Some insects are potential seed disperses, and other are seed predators. Other insect species eat plant flowers and leaves. We are investigating the nature of the species interactions in the alpine environment, where diversity is often lower than in forests. Manipulation experiments allow us to study the selective force on factors such as pollen limitation, outcrossing rate, floral attractiveness and species specificity.

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Date Range: 2007 - 2008

Funding Body: Massey University

Project Team:

Prof Mary Morgan-Richards - Project Leader
Prof Alastair Robertson - Team Member

Research Outputs

Journal

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Platania, L., Cardoso, A., Anderson, M., Fikáček, M., Gauthier, J., Hendrich, L., . . . Gómez-Zurita, J. (2024). New Caledonian rovers and the historical biogeography of a hyper-diverse endemic lineage of South Pacific leaf beetles. Systematic Entomology.
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Dowle, EJ., Trewick, SA., & Morgan-Richards, M. (2024). Fossil-calibrated phylogenies of Southern cave we̅tā show dispersal and extinction confound biogeographic signal. Royal Society Open Science. 11(2)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Shepherd, L., Simon, C., Langton-Myers, S., & Morgan-Richards, M. (2024). Insights into Aotearoa New Zealand’s biogeographic history provided by the study of natural hybrid zones. Journal of the Royal Society of New Zealand. 54(1), 55-74
[Journal article]Authored by: Morgan-Richards, M.

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Trewick, SA., Koot, EM., & Morgan-Richards, M. (2023). Māwhitiwhiti Aotearoa: Phylogeny and synonymy of the silent alpine grasshopper radiation of New Zealand (Orthoptera: Acrididae). Zootaxa. 5383(2), 225-241
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M. (2023). No barrier to fertilisation when different sexual populations of the mānuka stick insect are crossed. New Zealand Entomologist. 46(1-2), 1-6
[Journal article]Authored by: Morgan-Richards, M.

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Quenu, M., Judd, EJ., Morgan-richards, M., Trewick, SA., Holt, K., Tyler, J., . . . Lorrey, AM. (2023). High-resolution stable isotope profiles from shells of the land snail Placostylus reveal contrasting patterns between snails originating from New Zealand and New Caledonia. Journal of Quaternary Science. 38(7), 1171-1183
[Journal article]Authored by: Holt, K., Morgan-Richards, M., Trewick, S.

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Meza-Joya, FL., Morgan-Richards, M., Koot, EM., & Trewick, SA. (2023). Global warming leads to habitat loss and genetic erosion of alpine biodiversity. Journal of Biogeography. 50(5), 961-975
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Yarita, S., Morgan-Richards, M., & Trewick, SA. (2023). Genotypic detection of barriers to rat dispersal: Rattus rattus behind a peninsula predator-proof fence. Biological Invasions. 25(6), 1723-1738
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Marshall, CJ., Biggs, PJ., & Trewick, SA. (2023). Insect Freeze-Tolerance Downunder: The Microbial Connection. Insects. 14(1)
[Journal article]Authored by: Biggs, P., Morgan-Richards, M., Trewick, S.

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Nakano, M., Morgan-Richards, M., Clavijo-McCormick, A., & Trewick, S. (2023). Abundance and distribution of antennal sensilla on males and females of three sympatric species of alpine grasshopper (Orthoptera: Acrididae: Catantopinae) in Aotearoa New Zealand. Zoomorphology. 142(1), 51-62
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Godfrey, AJR., McKenzie, AO., & Morgan-Richards, M. (2023). Recommendations for non-lethal monitoring of tree wētā (Hemideina spp.) using artificial galleries. New Zealand Journal of Zoology. 50(3), 381-393
[Journal article]Authored by: Godfrey, A., Morgan-Richards, M.

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Carmelet-Rescan, D., Morgan-Richards, M., Pattabiraman, N., & Trewick, SA. (2022). Time-calibrated phylogeny and ecological niche models indicate Pliocene aridification drove intraspecific diversification of brushtail possums in Australia. Ecology and Evolution. 12(12)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Meza-Joya, FL., Morgan-Richards, M., & Trewick, SA. (2022). Relationships among body size components of three flightless New Zealand grasshopper species (Orthoptera, Acrididae) and their ecological applications. Journal of Orthoptera Research. 31(1), 91-103
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Cunha, AF., Carmelet-Rescan, D., Marques, AC., & Morgan-Richards, M. (2022). Contrasting morphological and genetic patterns suggest cryptic speciation and phenotype–environment covariation within three benthic marine hydrozoans. Marine Biology. 169(8)
[Journal article]Authored by: Morgan-Richards, M.

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Trewick, SA., Henderson, IM., Pohe, SR., & Morgan-Richards, M. (2022). Spatial Variation of Acanthophlebia cruentata (Ephemeroptera), a Mayfly Endemic to Te Ika-a-Māui—North Island of Aotearoa, New Zealand. Insects. 13(7)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Hegg, D., Morgan-Richards, M., & Trewick, SA. (2022). High alpine sorcerers: Revision of the cave wētā genus Pharmacus Pictet & de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies. European Journal of Taxonomy. 808, 1-58
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Koot, EM., Morgan-Richards, M., & Trewick, SA. (2022). Climate change and alpine-adapted insects: Modelling environmental envelopes of a grasshopper radiation. Royal Society Open Science. 9(3)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Nakano, M., Morgan-Richards, M., Trewick, SA., & Clavijo-McCormick, A. (2022). Chemical Ecology and Olfaction in Short-Horned Grasshoppers (Orthoptera: Acrididae). Journal of Chemical Ecology. 48(2), 121-140
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Pattabiraman, N., Morgan-Richards, M., Powlesland, R., & Trewick, SA. (2022). Unrestricted gene flow between two subspecies of translocated brushtail possums (Trichosurus vulpecula) in Aotearoa New Zealand. Biological Invasions. 24(1), 247-260
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Vilcot, M., & Trewick, SA. (2022). Lack of assortative mating might explain reduced phenotypic differentiation where two grasshopper species meet. Journal of Evolutionary Biology. 35(4), 509-519
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Quenu, M., Trewick, SA., Daly, EE., & Morgan-Richards, M. (2021). Generation of large mitochondrial and nuclear nucleotide sequences and phylogenetic analyses using high-throughput short-read datasets for endangered Placostylinae snails of the southwest Pacific. Molluscan Research. 41(3), 243-253
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Carmelet-Rescan, D., Morgan-Richards, M., Koot, EM., & Trewick, SA. (2021). Climate and ice in the last glacial maximum explain patterns of isolation by distance inferred for alpine grasshoppers. Insect Conservation and Diversity. 14(5), 568-581
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Brownstein, G., & Morgan-Richards, M. (2020). Indigenous plant naming and experimentation reveal a plant–insect relationship in New Zealand forests. Conservation Science and Practice. 2(10)
[Journal article]Authored by: Morgan-Richards, M.

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Trewick, SA., Taylor-Smith, B., & Morgan-Richards, M. (2021). Ecology and systematics of the wine wētā and allied species, with description of four new Hemiandrus species. New Zealand Journal of Zoology. 48(1), 47-80
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Daly, EE., Trewick, SA., Dowle, EJ., Crampton, JS., & Richards, MM. (2020). Conservation of pupu whakarongotaua the snail that listens for the war party. Ethnobiology and Conservation. 9
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Quenu, M., Trewick, SA., Brescia, F., & Morgan-Richards, M. (2020). Geometric morphometrics and machine learning challenge currently accepted species limits of the land snail Placostylus (Pulmonata: Bothriembryontidae) on the Isle of Pines, New Caledonia. Journal of Molluscan Studies. 86(1), 35-41
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Gemmell, MR., Hills, SFK., Marshall, BA., Beu, AG., Crampton, JS., . . . Morgan-Richards, M. (2020). Lineage identification affects estimates of evolutionary mode in marine snails. Systematic Biology. 69(6), 1106-1121
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Koot, EM., Morgan-Richards, M., & Trewick, SA. (2020). An alpine grasshopper radiation older than the mountains, on Kā Tiritiri o te Moana (Southern Alps) of Aotearoa (New Zealand). Molecular Phylogenetics and Evolution. 147
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Gemmell, MR., Trewick, SA., Hills, SFK., & Morgan-Richards, M. (2020). Phylogenetic topology and timing of New Zealand olive shells are consistent with punctuated equilibrium. Journal of Zoological Systematics and Evolutionary Research. 58(1), 209-220
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Taylor-Smith, B., Morgan-Richards, M., & Trewick, SA. (2020). Patterns of regional endemism among New Zealand invertebrates. New Zealand Journal of Zoology. 47(1), 1-19
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Hegg, D., Morgan-Richards, M., & Trewick, SA. (2019). Diversity and distribution of pleioplectron hutton cave wētā (Orthoptera: Rhaphidophoridae: Macropathinae), with the synonymy of weta chopard and the description of seven new species. European Journal of Taxonomy. 2019(577)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Langton-Myers, SS., & Trewick, SA. (2019). Loss and gain of sexual reproduction in the same stick insect. Molecular Ecology. 28(17), 3929-3941
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Nakano, M., Morgan-Richards, M., Godfrey, AJR., & McCormick, AC. (2019). Parthenogenetic females of the stick insect Clitarchus hookeri maintain sexual traits. Insects. 10(7)
[Journal article]Authored by: Godfrey, A., Morgan-Richards, M.

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Daly, EE., Walker, KJ., Morgan-Richards, M., & Trewick, SA. (2019). Spatial genetics of a high elevation lineage of Rhytididae land snails in New Zealand: the Powelliphanta Kawatiri complex. Molluscan Research. 39(3), 280-289
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Morgan-Richards, M., Daly, EE., & Trewick, SA. (2019). Tuatara and a new morphometric dataset for Rhynchocephalia: Comments on Herrera-Flores et al.. Palaeontology. 62(2), 321-334
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Gemmell, MR., Trewick, SA., Crampton, JS., Vaux, F., Hills, SFK., Daly, EE., . . . Morgan-Richards, M. (2018). Genetic structure and shell shape variation within a rocky shore whelk suggest both diverging and constraining selection with gene flow. Biological Journal of the Linnean Society. 125(4), 827-843
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Fitness, JL., Morgan-Richards, M., Hegg, D., & Trewick, SA. (2018). Reinstatement of the New Zealand cave wētā genus miotopus hutton (Orthoptera: Rhaphidophoridae) and description of a new species. European Journal of Taxonomy. 2018(468), 1-24
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Bulgarella, M., Quenu, M., Shepherd, LD., & Morgan-Richards, M. (2018). The ectoparasites of hybrid ducks in New Zealand (Mallard x Grey Duck). International Journal for Parasitology: Parasites and Wildlife. 7(3), 335-342
[Journal article]Authored by: Morgan-Richards, M.

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Sivyer, L., Morgan-Richards, M., Koot, E., & Trewick, SA. (2018). Anthropogenic cause of range shifts and gene flow between two grasshopper species revealed by environmental modelling, geometric morphometrics and population genetics. Insect Conservation and Diversity. 11(5), 415-434
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Mckean, NE., Trewick, SA., Griffin, MJ., Dowie, EJ., & Morgan-Rrichards, M. (2018). Viability and fertility of hybrid New Zealand tree we<sup>−</sup>ta<sup>−</sup> Hemideina spp. (Orthoptera: Anostostomatidae). Journal of Orthoptera Research. 27(2), 97-106
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Trewick, SA., Crampton, JS., Marshall, BA., Beu, AG., Hills, SFK., . . . Morgan-Richards, M. (2018). Evolutionary lineages of marine snails identified using molecular phylogenetics and geometric morphometric analysis of shells. Molecular Phylogenetics and Evolution. 127, 626-637
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Bridgeman, B., Morgan-Richards, M., Wheeler, D., & Trewick, SA. (2018). First detection of wolbachia in the New Zealand biota. PLoS ONE. 13(4)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Hills, SFK., Marshall, BA., Trewick, SA., & Morgan-Richards, M. (2018). Genome statistics and phylogenetic reconstructions for Southern Hemisphere whelks (Gastropoda: Buccinulidae). Data in Brief. 16, 172-181
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Bulgarella, M., Sivyer, L., Dowle, EJ., Hale, M., McKean, NE., . . . Trewick, SA. (2017). Explaining large mitochondrial sequence differences within a population sample. Royal Society Open Science. 4(11)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Monks, A., & Morgan-Richards, M. (2017). Male tree weta are attracted to cuticular scent cues but do not discriminate according to sex or among two closely related species. Ethology. 123(11), 825-834
[Journal article]Authored by: Morgan-Richards, M.

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Trewick, SA., Pilkington, S., Shepherd, LD., Gibb, GC., & Morgan-Richards, M. (2017). Closing the gap: Avian lineage splits at a young, narrow seaway imply a protracted history of mixed population response. Molecular Ecology. 26(20), 5752-5772
[Journal article]Authored by: Gibb, G., Morgan-Richards, M., Trewick, S.

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Bradley, E., Trewick, SA., & Morgan-Richards, M. (2017). Genetic distinctiveness of the Waikawa Island mouse population indicates low rate of dispersal from mainland New Zealand. NEW ZEALAND JOURNAL OF ECOLOGY. 41(2), 245-250
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Hills, SFK., Marshall, BA., Trewick, SA., & Morgan-Richards, M. (2017). A phylogeny of Southern Hemisphere whelks (Gastropoda: Buccinulidae) and concordance with the fossil record. Molecular Phylogenetics and Evolution. 114, 367-381
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Bradley, E., Trewick, SA., & Morgan-Richards, M. (2017). Genetic distinctiveness of the waikawa island mouse population indicates low rate of dispersal from Mainland New Zealand. New Zealand Journal of Ecology. 41(2)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Crampton, JS., Marshall, BA., Trewick, SA., & Morgan-Richards, M. (2017). Geometric morphometric analysis reveals that the shells of male and female siphon whelks Penion chathamensis are the same size and shape. Molluscan Research. 37(3), 194-201
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Trewick, SA., & Morgan-Richards, M. (2017). Speciation through the looking-glass. Biological Journal of the Linnean Society. 120(2), 480-488
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Taylor-Smith, BL., Trewick, SA., & Morgan-Richards, M. (2016). Three new ground wētā species and a redescription of Hemiandrus maculifrons. New Zealand Journal of Zoology. 43(4), 363-383
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Hills, SFK., Biggs, PJ., & Trewick, SA. (2016). Sticky genomes: Using NGS evidence to test hybrid speciation hypotheses. PLoS ONE. 11(5)
[Journal article]Authored by: Biggs, P., Hills, S., Morgan-Richards, M., Trewick, S.

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Nelson-Tunley, M., Morgan-Richards, M., & Trewick, SA. (2016). Genetic diversity and gene flow in a rare New Zealand skink despite fragmented habitat in a volcanic landscape. Biological Journal of the Linnean Society. 119(1), 37-51
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Mckean, NE., Trewick, SA., & Morgan-Richards, M. (2016). Little or no gene flow despite F<inf>1</inf> hybrids at two interspecific contact zones. Ecology and Evolution. 6(8), 2390-2404
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Vaux, F., Trewick, SA., & Morgan-Richards, M. (2016). Lineages, splits and divergence challenge whether the terms anagenesis and cladogenesis are necessary. Biological Journal of the Linnean Society. 117(2), 165-176
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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McKean, NE., Trewick, SA., & Morgan-Richards, M. (2015). Comparative cytogenetics of North Island tree we¯ta¯ in sympatry. New Zealand Journal of Zoology. 42(2), 73-84
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Dowle, EJ., Morgan-Richards, M., Brescia, F., & Trewick, SA. (2015). Correlation between shell phenotype and local environment suggests a role for natural selection in the evolution of Placostylus snails. Molecular Ecology. 24(16), 4205-4221
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Hinlo, AR., Smuts-Kennedy, C., Innes, J., Ji, W., Barry, M., . . . Hitchmough, RA. (2016). Identification of a Rare Gecko from North Island New Zealand, and Genetic Assessment of Its Probable Origin: A Novel Mainland Conservation Priority?. Journal of Herpetology. 50(1), 77-86
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M. (1997). Intraspecific karyotype variation is not concordant with allozyme variation in the Auckland tree weta of New Zealand, Hemideina thoracica (Orthoptera: Stenopelmatidae).. Biological journal of the Linnean Society. Linnean Society of London. 60(4), 423-442
[Journal article]Authored by: Morgan-Richards, M.

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Morgan‐richards, M., Daugherty, CH., & Gibbs, GW. (1995). Taxonomic status of tree weta from stephens island, mt holdsworth and mt arthur, based on allozyme variation. Journal of the Royal Society of New Zealand. 25(2), 301-312
[Journal article]Authored by: Morgan-Richards, M.

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Dowle, EJ., Morgan-Richards, M., & Trewick, SA. (2014). Morphological differentiation despite gene flow in an endangered grasshopper. BMC Evolutionary Biology. 14(1)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Bulgarella, M., Trewick, SA., Minards, NA., Jacobson, MJ., & Morgan-Richards, M. (2014). Shifting ranges of two tree weta species (Hemideina spp.): Competitive exclusion and changing climate. Journal of Biogeography. 41(3), 524-535
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Minards, NA., Trewick, SA., Godfrey, AJR., & Morgan-Richards, M. (2014). Convergent local adaptation in size and growth rate but not metabolic rate in a pair of parapatric Orthoptera species. Biological Journal of the Linnean Society.
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Goldberg, J., Morgan-Richards, M., & Trewick, SA. (2015). Intercontinental island hopping: Colonization and speciation of the grasshopper genus Phaulacridium (Orthoptera: Acrididae) in Australasia. Zoologischer Anzeiger. 255, 71-79
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Bulgarella, M., Trewick, SA., Godfrey, AJR., Sinclair, BJ., & Morgan-Richards, M. (2015). Elevational variation in adult body size and growth rate but not in metabolic rate in the tree weta Hemideina crassidens. Journal of Insect Physiology. 75, 30-38
[Journal article]Authored by: Godfrey, A., Morgan-Richards, M., Trewick, S.

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Mckean, NE., Trewick, SA., & Morgan-Richards, M. (2015). Comparative cytogenetics of North Island tree wētā in sympatry. New Zealand Journal of Zoology.
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Raubenheimer, D., & Morgan-Richards, M. (2013). Tolerance for nutrient imbalance in an intermittently feeding herbivorous cricket, the Wellington tree weta. PLoS ONE. 8(12)
[Journal article]Authored by: Morgan-Richards, M.

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Smith, BT., Morgan-Richards, M., & Trewick, S. (2013). New Zealand ground wētā (Anostostomatidae: Hemiandrus): Descriptions of two species with notes on their biology. New Zealand Journal of Zoology.
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

Myers, SS., Trewick, SA., & Morgan-Richards, M. (2013). Multiple lines of evidence suggest mosaic polyploidy in the hybrid parthenogenetic stick insect lineage Acanthoxyla. Insect Conservation and Diversity. 6(4), 537-548
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Dowle, EJ., Morgan-Richards, M., & Trewick, SA. (2013). Molecular evolution and the latitudinal biodiversity gradient. Heredity. 110(6), 501-510
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Jorgensen, M., & Morgan-Richards, M. (2013). Sex- and season-dependent behaviour in a flightless insect, the Auckland tree weta (Hemideina thoracica). New Zealand Journal of Ecology. 37(1)
[Journal article]Authored by: Morgan-Richards, M.

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Hills, SFK., Crampton, JS., Trewick, SA., & Morgan-Richards, M. (2012). DNA and Morphology Unite Two Species and 10 Million Year Old Fossils. PLoS ONE. 7(12)
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Fitness, J., Hitchmough, RA., & Morgan-Richards, M. (2012). Little and large: Body size and genetic clines in a New Zealand gecko (Woodworthia maculata) along a coastal transect. Ecology and Evolution. 2(2), 273-285
[Journal article]Authored by: Morgan-Richards, M.

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Chappell, EM., Trewick, SA., & Morgan-Richards, M. (2012). Shape and sound reveal genetic cohesion not speciation in the New Zealand orthopteran, Hemiandrus pallitarsis, despite high mitochondrial DNA divergence. Biological Journal of the Linnean Society. 105(1), 169-186
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Nakagawa, S., Trewick, SA., & Morgan-Richards, M. (2011). Does predation result in adult sex ratio skew in a sexually dimorphic insect genus?. Journal of Evolutionary Biology. 24(11), 2321-2328 Retrieved from http://onlinelibrary.wiley.com/doi/10.1111/jeb.2011.24.issue-11/issuetoc
[Journal article]Authored by: Morgan-Richards, M.

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Hills, SFK., Trewick, SA., & Morgan-Richards, M. (2011). Phylogenetic information of genes, illustrated with mitochondrial data from a genus of gastropod molluscs. BIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY. 104(4), 770-785
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Hills, SFK., Trewick, SA., & Morgan-Richards, M. (2011). Phylogenetic information of genes, illustrated with mitochondrial data from a genus of gastropod molluscs. Biological Journal of the Linnean Society. 104(4), 770-785
[Journal article]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

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Griffin, MJ., Morgan-Richards, M., & Trewick, SA. (2011). Is the tree weta Hemideina crassidens an obligate herbivore?. New Zealand Natural Sciences. 36, 11-19 Retrieved from https://sites.google.com/site/nznaturalsciences/home
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Nakagawa, S., Trewick, SA., & Morgan-Richards, M. (2011). Does predation result in adult sex ratio skew in a sexually dimorphic insect genus?. Journal of Evolutionary Biology. 24(11), 2321-2328
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., Wallis, GP., & Morgan-Richards, M. (2011). The invertebrate life of New Zealand: A phylogeographic approach. Insects. 2(3), 297-325
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wehi, PM., Nagakawa, S., Trewick, SA., & Morgan-Richards, M. (2011). Does predation result in adult sex ratio skew in a sexually dimorphic insect genus?. Journal of Evolutionary Biology.
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Griffin, MJ., Trewick, SA., Wehi, PM., & Morgan-Richards, M. (2011). Exploring the concept of niche convergence in a land without rodents: The case of weta as small mammals. New Zealand Journal of Ecology. 35(3)
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wyman, TE., Trewick, SA., Morgan-Richards, M., & Noble, ADL. (2011). Mutualism or opportunism? Tree fuchsia (Fuchsia excorticata) and tree weta (Hemideina) interactions. Austral Ecology. 36(3), 261-268
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Cook, LD., Trewick, SA., Morgan-Richards, M., & Johns, PM. (2010). Status of the New Zealand cave weta (Rhaphidophoridae) genera Pachyrhamma, Gymnoplectron and Turbottoplectron. Invertebrate Systematics. 24(2), 131-138
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Trewick, SA., & Stringer, IAN. (2010). Geographic parthenogenesis and the common tea-tree stick insect of New Zealand. Molecular Ecology. 19(6), 1227-1238
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Pratt, RC., Gibb, GC., Morgan-Richards, M., Phillips, MJ., Hendy, MD., & Penny, D. (2009). Toward resolving deep neoaves phylogeny: Data, signal enhancement, and priors. Molecular Biology and Evolution. 26(2), 313-326
[Journal article]Authored by: Gibb, G., Morgan-Richards, M.

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Trewick, SA., & Morgan Richards, M. (2009). New Zealand biology. Encyclopedia of Islands. , 665-673
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

Morgan-Richards, M., Smissen, RD., Shepherd, LD., Wallis, GP., Hayward, JJ., Chan, CH., . . . Chapman, HM. (2009). A review of genetic analyses of hybridisation in New Zealand. Journal of the Royal Society of New Zealand. 39(1), 15-34
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., Trewick, SA., Bartosch-Härlid, A., Kardailsky, O., Phillips, MJ., McLenachan, PA., . . . Penny, D. (2008). Bird evolution: Testing the Metaves clade with six new mitochondrial genomes. BMC Evolutionary Biology. 8(1)
[Journal article]Authored by: McLenachan, P., Morgan-Richards, M., Trewick, S.

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Trewick, SA., Morgan-Richards, M., & Collins, LJ. (2008). Are you my mother? Phylogenetic analysis reveals orphan hybrid stick insect genus is part of a monophyletic New Zealand clade. Molecular Phylogenetics and Evolution. 48(3), 799-808
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Pratt, RC., Morgan-Richards, M., & Trewick, SA. (2008). Diversification of New Zealand weta (Orthoptera: Ensifera: Anostostomatidae) and their relationships in Australasia. Philosophical Transactions of the Royal Society B: Biological Sciences. 363(1508), 3427-3437
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Trewick, SA., Bartosch-Haerlid, A., Kardailsky, O., Phillips, MJ., McLenachan, PA., . . . Penny, D. (2008). Bird evolution: testing the Metaves clade with six new mitochondrial genomes. BMC EVOLUTIONARY BIOLOGY. 8,
[Journal article]Authored by: McLenachan, P., Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Trewick, SA., & Dunavan, S. (2008). When is it coevolution? The case of ground wētā and fleshy fruits in New Zealand. New Zealand Journal of Ecology. 32(1), 108-112
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., & Trewick, SA. (2005). Hybrid origin of a parthenogenetic genus?. Molecular Ecology. 14(7), 2133-2142
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., Goldberg, J., & Morgan-Richards, M. (2005). Fewer species of Argosarchus and Clitarchus stick insects (Phasmida, Phasmatinae): Evidence from nuclear and mitochondrial DNA sequence data. Zoologica Scripta. 34(5), 483-491
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., & Morgan-Richards, M. (2005). After the deluge: Mitochondrial DNA indicates Miocene radiation and Pliocene adaptation of tree and giant weta (Orthoptera: Anostostomatidae). Journal of Biogeography. 32(2), 295-309
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., & Morgan Richards, M. (2004). Phylogenetics of New Zealand's tree, giant and tusked Weta (orthoptera: Anostostomatidae): Evidence from mitochondrial DNA. Journal of Orthoptera Research. 13(2), 185-196
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

Trewick, SA., Morgan-Richards, M., & Chapman, HM. (2004). Chloroplast DNA diversity of Hieracium pilosella (Asteraceae) introduced to New Zealand: Reticulation, hybridization, and invasion. American Journal of Botany. 91(1), 73-85
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., Trewick, SA., Chapman, HM., & Krahulcova, A. (2004). Interspecific hybridization among Hieracium species in New Zealand: Evidence from flow cytometry. Heredity. 93(1), 34-42
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., & Wallis, GP. (2003). A comparison of five hybrid zones of the weta Hemideina thoracica (Orthoptera: Anostostomatidae): Degree of cytogenetic differentiation fails to predict zone width. Evolution. 57(4), 849-861
[Journal article]Authored by: Morgan-Richards, M.

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Pinter, I., Bakker, F., Barrett, J., Cox, C., Gibby, M., Henderson, S., . . . Vogel, J. (2002). Phylogenetic and biosystematic relationships in four highly disjunct polyploid complexes in the subgenera Ceterach and Phyllitis in Asplenium (Aspleniaceae). Organisms Diversity and Evolution. 2(4), 299-311
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., Morgan-Richards, M., Russell, SJ., Henderson, S., Rumsey, FJ., Pintér, I., . . . Vogel, JC. (2002). Polyploidy, phylogeography and Pleistocene refugia of the rockfern Asplenium ceterach: Evidence from chloroplast DNA. Molecular Ecology. 11(10), 2003-2012
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M. (2002). Fission or fusion? Mitochondrial DNA phylogenetics of the chromosome races of Hemideina crassidens (Orthoptera: Anostostomatidae). Cytogenetic and Genome Research. 96(1-4), 217-222
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., & Gibbs, GW. (2001). A phylogenetic analysis of New Zealand giant and tree weta (Orthoptera : Anostostomatidae : Deinacrida and Hemideina) using morphological and genetic characters. Invertebrate Systematics. 15(1), 1-12
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., Trewick, SA., & Wallis, GP. (2001). Chromosome races with Pliocene origins: Evidence from mtDNA. Heredity. 86(3), 303-312
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Morgan-Richards, M., & Gibbs, GW. (2001). A phylogenetic analysis of New Zealand giant and tree weta (Orthoptera: Anostostomatidae: Deinacrida and Hemideina) using morphological and genetic characters. Invertebrate Taxonomy. 15(1), 1-12
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., Trewick, SA., & Wallis, GP. (2000). Characterization of a hybrid zone between two chromosomal races of the weta Hemideina thoracica following a geologically recent volcanic eruption. Heredity. 85(6), 586-592
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wallis, GP., Morgan-Richards, M., & Trewick, SA. (2000). Phylogeographical pattern correlates with Pliocene mountain building in the alpine scree weta (Orthoptera, Anostostomatidae). Molecular Ecology. 9(6), 657-666
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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105

Trewick, SA., & Morgan-Richards, M. (2000). Artificial weta roosts: A technique for ecological study and population monitoring of Tree Weta (Hemideina) and other invertebrates. New Zealand Journal of Ecology. 24(2), 201-208
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Wolff, K., Morgan-Richards, M., & Davison, AW. (2000). Patterns of molecular genetic variation in Plantago major and P. intermedia in relation to ozone resistance. New Phytologist. 145(3), 501-509
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M. (2000). Robertsonian translocations and B chromosomes in the Wellington tree weta, Hemideina crassidens (Orthoptera: Anostostomatidae). Hereditas. 132(1), 49-54
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., & Wolff, K. (1999). Genetic structure and differentiation of Plantago major reveals a pair of sympatric sister species. Molecular Ecology. 8(6), 1027-1036
[Journal article]Authored by: Morgan-Richards, M.

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Wolff, K., & Morgan-Richards, M. (1998). PCR markers distinguish Plantago major subspecies. Theoretical and Applied Genetics. 96(2), 282-286
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M. (1997). Intraspecific karyotype variation is not concordant with allozyme variation in the Auckland tree weta of New Zealand, Hemideina thoracica (Orthoptera: Stenopelmatidae). Biological Journal of the Linnean Society. 60(4), 423-442
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., & Gibbs, GW. (1996). Colour, allozyme and karyotype variation show little concordance in the New Zealand Giant Scree Weta Deinacrida connectens (Orthoptera: Stenopelmatidae). Hereditas. 125(2-3), 265-276
[Journal article]Authored by: Morgan-Richards, M.

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Morgan-Richards, M., & Townsend, JA. (1995). Hybridisation of tree weta on banks peninsula, new zealand, and colour polymorphism within hemideina ricta (Orthoptera: Stenopelmatidae). New Zealand Journal of Zoology. 22(4), 393-399
[Journal article]Authored by: Morgan-Richards, M.

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Trewick, SA., & Trewick, SA. (1995). On the distribution of tree weta in the North Island, New Zealand. Journal of the Royal Society of New Zealand. 25(4), 485-493
[Journal article]Authored by: Morgan-Richards, M., Trewick, S.

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Book

Trewick, SA., & Morgan-Richards, M. (2016). Phylogenetics and conservation in New Zealand: The long and the short of it. In R. Pellens, & P. Grandcolas (Eds.) Biodiversity Conservation and Phylogenetic Systematics: Preserving our evolutionary heritage in an extinction crisis. (pp. 81 - 98). : Springer International
[Chapter]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, S., & Morgan-Richards, M.(2014). NZ Wild Life Introducing the Weird and Wonderful Character of Natural New Zealand. : Penguin Books
[Authored Book]Authored by: Morgan-Richards, M., Trewick, S.

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Trewick, SA., & Morgan-Richards, M.(2005). Stick insects. Auckland, NZ: Reed Publishing
[Authored Book]Authored by: Morgan-Richards, M., Trewick, S.

Morgan Richards, M., King, T., & Trewick, SA. (2001). The evolutionary history of Tree Weta: A genetic approach. In LHF. Ed (Ed.) The Biology of Wetas, King Crickets and their Allies. (pp. 111 - 126). Oxon, UK: CABI Publishing
[Chapter]Authored by: Morgan-Richards, M., Trewick, S.

Conference

Hills, S., Vaux, F., Trewick, S., Crampton, J., Marshall, B., Beu, A., . . . Morgan-Richards, M. (2018). Identifying evolutionary lineages of marine snails.. Poster session presented at the meeting of Molluscs 2018, the triennial meeting of the Malacological Society of Australasia.. Wellington, New Zealand
[Conference Poster]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Hills, S., Gemmell, M., Crampton, J., Trewick, S., & Morgan-Richards, M. (2018). Do New Zealand Olive Shells (Amalda spp.) support Punctuated Equilibria?. Poster session presented at the meeting of Molluscs 2018, the triennial meeting of the Malacological Society of Australasia.. Wellington, New Zealand
[Conference Poster]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

King, S., Barry, M., Morgan-Richards, M., & Trewick, S. (2017). The biology and ecology of green gecko (Naultinus) found in unique habitat on the Denniston and Stockton Plateaus. Poster session presented at the meeting of 2017 New Zealand Molecular Ecology Conference. The Catlins, New Zealand
[Conference Poster]Authored by: Morgan-Richards, M., Trewick, S.

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Hills, SFK., Gemmell, M., Vaux, F., Marshall, BA., Crampton, JS., Trewick, SA., . . . Morgan-Richards, M. (2015, November). Mitochondrial Phylogenomics of New Zealand Aeneator Finlay, 1926 and Buccinulum Deshayes, 1830 species (Buccinidae).. Presented at Molluscs 2015, the triennial meeting of the Malacological Society of Australasia.. Coffs Harbour, Australia.
[Conference Oral Presentation]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Hills, SFK., Crampton, JS., Trewick, SA., & Morgan-Richards, . (2014, February). Diversification patterns in molecular and paleontological data of the marine mollusc genus Alcithoe.. Presented at Waiheke 2014, the 18th Annual New Zealand Phylogenomics Meeting. Waiheke Island, New Zealand.
[Conference Oral Presentation]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Hills, SFK., Crampton, JS., Marshall, BA., Beu, AG., Trewick, SA., & Morgan-Richards, M. (2014). Evolution of the New Zealand Alcithoe, an integration of molecular and paleontological evidence.. In Abstracts: ‘GeoGenes V’ Geology and Genes V 2014 – a meeting of mudstone and molecules, ‘GeoGenes V’ Geology and Genes V 2014 – a meeting of mudstone and molecules Wellington, New Zealand: Geoscience Society of New Zealand
[Conference Abstract]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Hills, SFK., Crampton, JS., Trewick, SA., & Morgan-Richards, M. (2013). Fossils and phylogeny: comparing patterns of extinction and speciation.. In AÇOREANA, Revista de Estudos Açoreanos Vol. Suplemento 8, Julho de 2013 , World Congress of Malacology 2013 Ponta Delgada: Sociedade Afonso Chaves
[Conference Abstract]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Hills, SFK., Crampton, JS., Trewick, SA., & Morgan-Richards, M. (2012, December). The evolution of New Zealand Alcithoe, an integration of molecular and paleontological evidence.. Presented at Molluscs 2012, the triennial meeting of the Malacological Society of Australasia. Melbourne, Australia.
[Conference Oral Presentation]Authored by: Hills, S., Morgan-Richards, M., Trewick, S.

Morgan-Richards, M., Fitness, JL., Ball, OJP., Godfrey, AJR., & Trewick, SA. (2014). Species ecology - the distribution and abundance of forest invertebrates. Poster session presented at the meeting of The New Zealand Ecological Society Conference, Palmerston North, New Zealand
[Conference Poster]Authored by: Godfrey, A., Morgan-Richards, M., Trewick, S.

Godfrey, AJR., Bulgarella, M., Trewick, SA., & Morgan-Richards, M. (2014, November). Modelling growth of captive tree weta. Presented at New Zealand Ecological Society Conference. Palmerston North, New Zealand.
[Conference Oral Presentation]Authored by: Godfrey, A., Morgan-Richards, M., Trewick, S.

Godfrey, AJR., Bulgarella, M., Trewick, SA., & Morgan-Richards, M. (2014, November). Modelling growth of captive tree weta. Presented at 65th Annual Conference of the New Zealand Statistical Association. Wellington, New Zealand.
[Conference Oral Presentation]Authored by: Godfrey, A., Morgan-Richards, M., Trewick, S.

Lehnebach, C., Frericks, J., Zeller, A., Keyzers, R., Ong, CP., Morgan-Richards, M., . . . Robertson, AW. (2013, December). Spider orchids and fungus gnats: a story about deception, sex and death in New Zealand forests. Presented at 5th Internationalk Orchid Conservation Congress. Saint-Denis de la Réunion.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Robertson, A.

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Morgan-Richards, M., Bulgarella, M., Minards, N., & Trewick, SA. (2013, January). Range shifts and adaptation to local conditions in parapatric insects. Presented at VII Southern Connections Congress 2013. Dunedin New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Trewick, SA., & Morgan-Richards, M. (2011, August). Hopping across the ditch: The New Zealand Orthoptera. Presented at The 3rd Combined Australian and New Zealand Entomological Societies Conference. Lincoln University, Christchurch, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Fitness, J., Trewick, SA., Morgan-Richards, M., & Cook, L. (2011). Beta Weta Geta: Rhaphidophoridae taxonomy in New Zealand. Poster session presented at the meeting of The 3rd Combined Australian and New Zealand Entomological Societies Conference. Lincoln University, Christchurch, New Zealand
[Conference Poster]Authored by: Morgan-Richards, M., Trewick, S.

Trewick, SA., Morgan-Richards, M., Goldberg, J., Pratt, R., & Johns, P. (2010, February). Better the weta you know: biogeography of southern orthoptera. Presented at Southern Connection Congress. Bariloche, Argentina.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Griffin, M., Morgan-Richards, M., & Trewick, SA. (2010, November). Are tree weta (Hemideina crassidens) herbivores?. Presented at Biodiversity: 2010 and beyond. New Zealand Ecological Society Conference. University of Otago, Dunedin, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Minards, NA., Trewick, SA., & Morgan-Richards, M. (2010, November). Competitive exclusion and cold adaptation of two tree weta species. Presented at Biodiversity: 2010 and beyond. New Zealand Ecological Society Conference. University of Otago, Dunedin, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Wehi, PM., Jorgensen, M., & Morgan-Richards, M. (2010, November). Sex- and season-dependent behaviour in the Auckland tree weta. Presented at Biodiversity: 2010 and beyond. New Zealand Ecological Society Conference. University of Otago, Dunedin, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Morgan-Richards, M., Jacobson, M., Minards, NA., & Trewick, SA. (2010, November). Understanding climate change: tree weta as model organisims. Presented at Biodiversity: 2010 and beyond. New Zealand ecological Society Conference. University of Otago, Dunedin, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Wehi, PM., Jorgensen, M., & Morgan-Richards, M. (2010, November). Sex- and seasonal-dependent behaviour in the Auckland tree weta. Presented at New Zealand Ecological Society Annual Conference. Dunedin, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Myers, S., & Morgan-Richards, M. (2009, November). Investigating hybrid origins of Acanthoxyla, New Zealand's parthenogenetic genus, using combined genetic techniques. Presented at Molecular Ecology annual meeting. Tautuku, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Fitness, J., Hitchmough, RA., & Morgan-Richards, M. (2009, November). Wellington geckos meet Wairarapa geckos; genetic cline within Hoplodactylus maculatus. Presented at Molecular Ecology annual meeting. Tautuku, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Myers, SS., & Morgan Richards, M. (2009, November). Investigating hybrid origins of Acanthoxyla New Zealand's parthenogenetic genus using combined genetic techniques. Presented at Annual Meeting MolEcol09. Tautuku, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

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Fitness, JL., Hitchmough, RA., & Morgan Richards, M. (2009, November). Wellington geckos meet Wairarapa geckos; genetic cline within Hoplodactylus maculatus. Presented at Annual Meeting MolEcol09. Tautuku, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

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Morgan Richards, M., Trewick, SA., & Welvaert, S. (2009, November). Tree weta phylogeography and the trouble of paralogs. Presented at Annual Meeting MolEco109. Tautuku, New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

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Chappell, E., Trewick, S., Morgan-Richards, M., & Castro, IC. (2008). Morphology, phylogeography and drumming behaviour in a New Zealand Ground Weta, Hemiandrus pallitarsis. Presented at New Zealand Ecological Society Conference. Auckland.
[Conference Oral Presentation]Authored by: Castro, I., Morgan-Richards, M., Trewick, S.

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Morgan Richards, M., Hills, SF., Marshall, B., & Crampton, JS. (2008, November). Rapid morphological evolution or mtDNA introgression?. Presented at 11th Annual Molecular Ecology Meeting. Lake Okataina, Rotorua, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Morgan Richards, M., Hills, SF., Marshall, B., & Crampton, JS. (2008). Reconciling morphological divergence and genetic differentation: two examples. In Proceedings of the Speciation in Molluscs, NHM Meeting(pp. 10). : The Malacologiest: The Bulletin of the Malacological Society of London
[Conference Abstract]Authored by: Hills, S., Morgan-Richards, M.

Pratt, RC., Morgan-Richards, M., & Trewick, SA. (2007, June). Witness to the sinking ark or victims of the Pacific flypaper? Phylogenetic relationships of Anostostomatid crickets.. Presented at Evolution 2007. Christchurch Convention Centre, Christchurch, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Morgan-Richards, M., & Trewick, SA. (2007, June). New Zealand stick insects. Presented at Evolution 2007 Joint meeting of the society for the study of evolution, American Society of Naturalists and Systemtic Biologists. Christchurch New Zealand.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Cook, L., Trewick, SA., & Morgan Richards, M. (2007, December). Gene flow and population genetics of the New Zealand cave weta. Presented at 10th Annual New Zealand Molecular Ecology Meeting. Kaikoura, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Chappell, E., Trewick, SA., Morgan Richards, M., & Castro, IC. (2007, December). Phylogeography, morphology and reproductive behaviour of a New Zealand ground weta Hemiandrus pallitarsis. Presented at 10th Annual New Zealand Molecular Ecology Meeting. Kaikoura, NZ.
[Conference Oral Presentation]Authored by: Castro, I., Morgan-Richards, M., Trewick, S.

Morgan Richards, M. (2007, December). Geographic parthenogenesis in New Zealand stick insects. Presented at 10th Annual New Zealand Molecular Ecology Meeting. Kaikoura, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Chappell, E., Trewick, SA., Morgan Richards, M., & Castro, IC. (2007, November). Phylogeography, morphology and reproductive behaviour of ground weta (Hemiandrus pallitarsis). Presented at New Zealand Ecological Society Conference. University of Canterbury, Christchurch, NZ.
[Conference Oral Presentation]Authored by: Castro, I., Morgan-Richards, M., Trewick, S.

Morgan-Richards, M. (2006). Does gene flow limit species? Distinguishing between reproductive assurance and out-breeding depression avoidance.. Poster session presented at the meeting of Genetics of Speciation Conference. Vancouver Canada
[Conference Poster]Authored by: Morgan-Richards, M.

Morgan Richards, M., & Trewick, SA. (2005, June). Hybrid origins of a parthenogenetic genus?. Presented at Molecular Biology & Evolution 2005. Auckland, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Morgan Richards, M., & Trewick, SA. (2005, April). Hybrid origins of a parthenogenetic genus?. Presented at 54th Annual Conference of the Entomological Society of New Zealand. Napier, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Pratt, RC., Morgan Richards, M., & Trewick, SA. (2005). Anostostomatid phylogenetics: What DNA can tell us. Poster session presented at the meeting of 54th Annual Conference of the Entomological Society of New Zealand. Napier, NZ
[Conference Poster]Authored by: Morgan-Richards, M., Trewick, S.

Pratt, RC., Morgan Richards, M., & Trewick, SA. (2005). Weta worldwide: Systematics and biogeography of the Anostostomatids. Poster session presented at the meeting of Molecular Biology & Evolution 2005. Auckland, NZ
[Conference Poster]Authored by: Morgan-Richards, M., Trewick, S.

Pratt, RC., Morgan Richards, M., & Trewick, SA. (2004, December). Evolutionary relationships within Anostostomatidae. Presented at 7th New Zealand Molecular Ecology Conference. Pohangina, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Morgan Richards, M. (2003, December). Chromosome and genome evolution of New Zealand's stick insects. Presented at Molecular Ecology Annual Meeting: MolEcol03. Pounawea, Southland, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Morgan Richards, M., Trewick, SA., & Vogal, JC. (2001, November). European fern phylogeography. Presented at 4th New Zealand Molecular Ecology Conference. Rotoiti Lodge Outdoor Education Centre, St. Arnaud, Nelson Lakes, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Morgan Richards, M. (2001, January). Can relative hybrid zone width be predicted from number of chromosome rearrangements?: A comparison of five zones of the weta Hemideina thoracica (Orthoptera: Anostostomatidae). Presented at 34th Annual Meeting of the Population Genetics Group. University of Sheffield, Sheffield, UK.
[Conference Oral Presentation]Authored by: Morgan-Richards, M.

Morgan Richards, M., Trewick, SA., & Vogal, JC. (2001, November). European fern phylogeography. Presented at 4th New Zealand Molecular Ecology Conference. Nelson, NZ.
[Conference Oral Presentation]Authored by: Morgan-Richards, M., Trewick, S.

Wolff, K., & Morgan-Richards, M.(1997, August). The use of RAPD data in the analysis of population genetic structure: case studies of Alkanna (Boraginaceae) and Plantago (Plantaginaceae). MOLECULAR SYSTEMATICS AND PLANT EVOLUTION. (pp. 51 - 73). 0309-2593.
[Conference]Authored by: Morgan-Richards, M.

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Teaching and Supervision

Teaching Statement

Evolution [196.207]

Ecology & Conservation [196.205]

Animal Biodiversity [199.317]

New Zealand Environments [121.103]

Biology of Animals [199.101]

Entomology [199.717]

Conservation Biology [232.701]

Topics in Biodiveristy [199.718]

Contact Supervisor

Summary of Doctoral Supervision

Position	Current	Completed
Main Supervisor	1	6
Co-supervisor	3	6

Current Doctoral Supervision

Main Supervisor of:

Nyasha Chikwature - Doctor of Philosophy
Evaluating ecologically sustainable ways to disrupt the Hemiandrus bilobatus - grape vine association

Co-supervisor of:

Cheten Dorji - Doctor of Philosophy
Conservation of Himalayan crickets (Orthoptera: Rhaphidophoridae) in the context of anthropogenic climate change
Leonardo Meza Joya - Doctor of Philosophy
Global warming responses within the New Zealand alpine radiation of acridid grasshoppers
Nim Pattabiraman - Doctor of Philosophy
Population Genetics and Genomics of a Marsupial Species: Analysis of Native and Invasive Brushtail Possum Populations (Trichosurus vulpecula)

Completed Doctoral Supervision

Main Supervisor of:

2024 - Mari Nakano - Doctor of Philosophy
Food plants and chemical ecology of sympatric species of endemic New Zealand alpine grasshoppers
2021 - Mathieu Quenu - Doctor of Philosophy
Evolution and Stable Isotopes in Placostylus species of the southwest Pacific
2017 - Felix Vaux - Doctor of Philosophy
Evolutionary lineages and the Diversity of New Zealand true whelks
2017 - Michael Gemmell - Doctor of Philosophy
Genetic and Phenotypic Lineages in Neogastropod Molluscs: A Journey Through Time and Morphospace
2017 - Elizabeth Daly - Doctor of Philosophy
Fine scale population structure through space and time.
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2010 - Simon Hills - Doctor of Philosophy
The evolution of a marine gastropod genus: Rocks, clocks and convergence
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Co-supervisor of:

2023 - David Carmelet-Rescan - Doctor of Philosophy
Genomic differentiation of brushtail possum (Trichosurus vulpecula) populations
2018 - Emily Koot - Doctor of Philosophy
The Ecology and Evolution of New Zealand’s Endemic Alpine Grasshoppers
2017 - Josephine Fitness - Doctor of Philosophy
Te Putaiao o Tokoriro: Taxonomy and diversity of New Zealand cave weta (Orthoptera: Rhapidophoridae)
2015 - Briar Smith - Doctor of Philosophy
Evolution of diversity: Analysis of species and speciation in Hemiandrus ground weta
2014 - Edwina Dowle - Doctor of Philosophy
Rates of Molecular Evolution and Gene Flow
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2011 - Julia Goldberg - Doctor of Philosophy
Speciation and phylogeography in the New Zealand archipelago
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Media and Links

Media

20 Aug 2019 - Radio, Online
UK news coverge of original research
The stick insect population that lost its sex drive
19 Aug 2019 - Newspaper
UK media coverage
No sex please, we're British (stick insects) This article is more than 1 year old Phasmids hailing from New Zealand become asexual after arriving in the UK
05 Aug 2021 - Newspaper
Coverage of new research
Two grasshopper species interbreeding

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Prof Mary Morgan-Richards PhD

Professor in Evolutionary Biology

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Research Interests

Research Opportunities

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Area of Expertise

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Research Projects

Summary of Research Projects

Completed Projects

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Conference

Teaching and Supervision

Teaching Statement

Summary of Doctoral Supervision

Current Doctoral Supervision

Completed Doctoral Supervision

Media and Links

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